[R] About "duleg" Indicator Species Analysis

[Mariko Takemoto]

I have a question about the function "duleg" in the
package "labdsv".
How the p-value is calculated ?
Dufrene and Legendre (1997) have shown two calculation way
; one is the difference between the observed value and the
mean of those obtained from random permutations and
another is the rank of the observed value in the randomly
generated distribution ordered in decreasing order.
Which way does this function treat?