[R] Problem with "different" proj4string that are actually identical

[Hemery, Lenaig]

Dear everybody
I am trying to run an Ecological Niche Factor Analysis (enfa) with the package "adehabitatHS" but before I do so, I want to test the significance of its parameters marginality and tolerance, by using the function niche.test() that runs a Monte Carlo test. I used to do that with the old package "adehabitat" that is now deprecated so I updated all my codes. Everything I want to do seems to work fine except the niche.test() function and I get the error message here below.

> testCro<-niche.test(paramdf, spCro, nrep = 999, o.include = TRUE)
Error in join(pts, ta) : different proj4string in x and xy

I checked the proj4string for x (paramdf) and xy (spCro) and there are exactly identical (see below). 

> summary(spCro)
Object of class SpatialPoints
Coordinates:
              min     max
longitude -124.91 -124.13
latitude    39.86   46.95
Is projected: FALSE
proj4string :
[+proj=longlat +datum=WGS84 +ellps=WGS84 +towgs84=0,0,0]
Number of points: 84

> summary(paramdf)
Object of class SpatialPixelsDataFrame
Coordinates:
         min       max
x -125.50000 -123.7998
y   39.50384   47.0000
Is projected: FALSE
proj4string :
[+proj=longlat +datum=WGS84 +ellps=WGS84 +towgs84=0,0,0]
Number of points: 108237
Grid attributes:
  cellcentre.offset cellsize cells.dim
x        -125.49517  0.00966       176
y          39.50867  0.00966       776
Data attributes:
      bath              slope              rugo               sumT      
 Min.   :-3611.83   Min.   : 0.0000   Min.   :0.000000   Min.   : 1.184 
 1st Qu.:-1885.11   1st Qu.: 0.5271   1st Qu.:0.008044   1st Qu.: 2.150 
 Median : -894.61   Median : 1.4974   Median :0.036921   Median : 3.978 
 Mean   :-1157.85   Mean   : 3.4237   Mean   :0.147369   Mean   : 4.794 
 3rd Qu.: -150.85   3rd Qu.: 4.1152   3rd Qu.:0.197974   3rd Qu.: 7.388 
 Max.   :   -0.05   Max.   :61.3592   Max.   :0.992856   Max.   :14.436 
[...]

I think I looked for all the help I could find on the internet and didn't find any solution so here is my last chance.
Does anybody have an idea on how to solve this problem?
By the way, I use the R version 3.1.3 and up-to-date "adehabitatHS", "ade4", "sp" and other packages.
Thanks a lot,
Lenaig

≈≈≈≈ ><)))°> <°(((>< ≈≈≈≈
Lenaïg Hemery, Ph. D.
Research Associate - post-doc
Hatfield Marine Science Center
Oregon State University
2030 S.E. Marine Science Drive, Newport, OR 97365, USA
Cell:+1-541-272-7196